Quantitative understanding of biological systems through the application of physical principles. Course will emphasize topics that span multiple length and time scales, and different levels of biological organization. Two to four topics per semester, including possibly organismal motion from molecular processes to whole organisms, nervous systems from membrane channels to neuronal networks, noise in biology, novel biophysical technologies, etc. Recommended prerequisite: Biology 201L or 203L, Mathematics 212 and 216 or equivalent, and calculus-based introductory physics or permission of the instructors.